Whole wheat flour contains bran and germ, which interfere with gluten development, resulting in a denser loaf compared to bread made with all-purpose flour. All-purpose flour is primarily endosperm, allowing for better gluten formation and a lighter texture.
The density difference between whole wheat bread and all-purpose bread stems from the composition of the flours and their impact on gluten development. Gluten, a protein complex formed when wheat flour is mixed with water, provides the structure and elasticity necessary for bread to rise.
All-purpose flour is milled from the endosperm of the wheat kernel, which is primarily starch and protein. This allows for relatively unimpeded gluten development, resulting in a lighter, airier texture.
Whole wheat flour, on the other hand, includes the bran and germ of the wheat kernel in addition to the endosperm. The bran, with its sharp edges, physically cuts and disrupts the gluten strands as they form. The germ contains oils that also interfere with gluten development. This disruption prevents the gluten from forming a strong, elastic network, leading to a denser, less airy loaf. The presence of more fiber in whole wheat flour also absorbs more water, which can contribute to a heavier texture.
To improve the texture of whole wheat bread, try using a technique called an autolyse. This involves mixing the flour and water together and letting it rest for 20-30 minutes before adding the yeast and other ingredients. This allows the flour to fully hydrate and the gluten to begin developing, resulting in a slightly lighter and more open crumb.