Whole wheat flour contains bran and germ, which interfere with gluten development, resulting in a denser loaf of bread compared to bread made with all-purpose flour. All-purpose flour is primarily endosperm, allowing for better gluten formation and a lighter texture.
The density difference between white bread made with whole wheat flour and all-purpose flour stems from the composition of each flour and its impact on gluten development. Gluten, a protein complex formed when wheat flour is mixed with water, provides the structure and elasticity necessary for a light and airy bread.
All-purpose flour is primarily composed of the endosperm of the wheat kernel. The endosperm is mostly starch and protein, which are ideal for gluten formation. When all-purpose flour is hydrated and kneaded, the gluten proteins align and create a network that traps gases produced by yeast, causing the dough to rise.
Whole wheat flour, on the other hand, includes the bran and germ of the wheat kernel in addition to the endosperm. The bran, with its sharp edges, physically cuts and disrupts the gluten strands, preventing them from forming a strong, cohesive network. The germ contains oils that can also interfere with gluten development. This disruption results in a weaker gluten structure, less gas retention, and ultimately, a denser loaf of bread. The higher fiber content in whole wheat flour also absorbs more water, which can further contribute to a denser texture.
To improve the texture of whole wheat bread, try using a technique called an autolyse. This involves mixing the flour and water and letting it rest for 20-30 minutes before adding the yeast and salt. This allows the flour to fully hydrate, promoting better gluten development despite the presence of bran.