Whole wheat flour contains bran and germ, which interfere with gluten development, resulting in a denser loaf compared to bread made with all-purpose flour. All-purpose flour is primarily endosperm, allowing for better gluten formation and a lighter texture.
The density difference between white bread made with whole wheat flour and all-purpose flour stems from the composition of each flour and its impact on gluten development. Gluten, a protein complex formed when wheat flour is mixed with water, provides the structure and elasticity necessary for a light and airy bread.
All-purpose flour is milled from the endosperm of the wheat kernel, the starchy part that is low in protein and fiber. This allows for optimal gluten development, creating a strong network that traps gases produced during fermentation, resulting in a light and airy texture.
Whole wheat flour, on the other hand, includes the bran and germ of the wheat kernel in addition to the endosperm. The bran, with its sharp edges, physically interferes with gluten formation, cutting the gluten strands and preventing them from forming a strong, cohesive network. The germ contains oils that can also inhibit gluten development. This weaker gluten structure is less able to trap gases, leading to a denser, less voluminous loaf.
Furthermore, whole wheat flour absorbs more water than all-purpose flour. This higher water absorption can make the dough heavier and more difficult to work with, further contributing to the denser texture of the final product.
To improve the texture of whole wheat bread, try using a technique called an autolyse. This involves mixing the flour and water together and letting it rest for 20-30 minutes before adding the yeast and other ingredients. This allows the flour to fully hydrate and the gluten to begin developing, resulting in a slightly lighter and more open crumb.