Whole wheat flour contains bran, which interferes with gluten development, resulting in a denser brioche compared to using all-purpose flour. All-purpose flour, being refined, allows for better gluten formation and a lighter, airier texture.
The key difference lies in the composition of the flours. All-purpose flour is milled from the endosperm of the wheat kernel, resulting in a fine, consistent powder that readily forms gluten when hydrated and kneaded. Gluten is the protein network that gives bread its structure and elasticity, allowing it to rise and hold its shape.
Whole wheat flour, on the other hand, includes the bran and germ of the wheat kernel in addition to the endosperm. The bran, being sharp and fibrous, physically cuts and disrupts the gluten strands as they form. This interference weakens the gluten network, preventing it from trapping as much air during fermentation. Consequently, the brioche made with whole wheat flour will have a tighter crumb and a denser texture. The germ also contains oils that can interfere with gluten development.
Brioche, in particular, relies heavily on strong gluten development to support its rich, buttery dough. The added fat in brioche can already inhibit gluten formation to some extent, so using whole wheat flour exacerbates this issue, leading to a significantly denser final product.
If you want to incorporate whole wheat flour into your brioche, try substituting only a portion of the all-purpose flour (e.g., 25-50%) with whole wheat. You can also try autolysing the flour (mixing the flour and water and letting it rest for 20-30 minutes) before adding the other ingredients to improve gluten development.