Whole wheat flour contains bran and germ, which interfere with gluten development, resulting in a denser brioche compared to using all-purpose flour. All-purpose flour, being mostly endosperm, allows for better gluten formation and a lighter, airier texture.
The density difference between brioche made with whole wheat flour and all-purpose flour stems from the composition of each flour and its impact on gluten development. Gluten, formed from the proteins glutenin and gliadin when flour is hydrated and kneaded, provides the structure and elasticity necessary for a light and airy brioche.
All-purpose flour is primarily composed of the endosperm of the wheat kernel. The endosperm is mostly starch and protein, allowing for strong gluten development when mixed with water and kneaded. This strong gluten network traps the gases produced by the yeast during fermentation, resulting in a light and airy texture.
Whole wheat flour, on the other hand, includes the bran and germ of the wheat kernel in addition to the endosperm. The bran, with its sharp edges, physically interferes with gluten formation, cutting the gluten strands and preventing them from forming a strong, cohesive network. The germ contains oils that can also inhibit gluten development. As a result, the gluten network in whole wheat brioche is weaker and less able to trap gases effectively, leading to a denser, less airy final product. The higher fiber content in whole wheat flour also absorbs more liquid, which can further contribute to a denser texture.
If you want to use whole wheat flour in brioche, try substituting only a portion of the all-purpose flour with whole wheat (e.g., 25-50%). This will add some of the nutritional benefits and flavor of whole wheat without significantly compromising the texture. You may also need to increase the hydration slightly to compensate for the higher water absorption of whole wheat flour.